[1] NOAA, Alaska Fisheries Sci Ctr, Seattle, WA USA
[2] Oregon State Univ, Coll Ocean & Atmospher Sci, Corvallis, OR 97331 USA
[3] Tech Univ Denmark, Natl Inst Aquat Resources, Charlottenlund, Denmark
[4] Univ Hamburg, Inst Hydrobiol & Fisheries Sci, Hamburg, Germany
[5] McGill Univ, Dept Biol, Montreal, PQ H3A 1B1, Canada
[6] Inst Marine Res, N-5024 Bergen, Norway