[1] Univ Tokyo, Grad Sch Arts & Sci, Dept Gen Syst Studies, Tokyo 1538902, Japan
[2] Nagasaki Univ, Inst Trop Med, Dept Int Hlth, Nagasaki 8528523, Japan
[3] Univ Basel, Dept Environm Sci, Zool, CH-4051 Basel, Switzerland
[4] Kyushu Univ, Fac Engn Sci, Interdisciplinary Grad Sch Engn Sci, Dept Energy & Environm Engn, Fukuoka, Fukuoka 8168580, Japan
[5] Kyushu Univ, Fac Engn Sci, Dept Adv Environm Sci & Engn, Fukuoka, Fukuoka 8168580, Japan