[1] Univ Guelph, Dept Environm Biol, Guelph, ON N1G 2W1, Canada
[2] Georgia So Univ, Inst Arthropodol & Parasitol, Statesboro, GA 30460 USA
[3] Univ Toronto, Dept Zool, Toronto, ON M5S 3G5, Canada
[4] USDA ARS, Biosystemat & Natl Parasite Collect Unit, BARC E 1180, Beltsville, MD 20715 USA