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Gansu Agr Univ, Coll Grassland Sci, Key Lab Grassland Ecosyst, Minist Educ,Engn & Technol Res Ctr Alpine Rodent, Lanzhou 730070, Peoples R China
Ningxia Acad Agr & Forestry Sci, Guyuan Branch, Guyuan 756000, Peoples R China Gansu Agr Univ, Coll Grassland Sci, Key Lab Grassland Ecosyst, Minist Educ,Engn & Technol Res Ctr Alpine Rodent, Lanzhou 730070, Peoples R China