[1] Royal Melbourne Hosp, Stem Cells & Canc Div, Melbourne, Vic, Australia
[2] Walter & Eliza Hall Inst Med Res, Stem Cells & Canc Div, Melbourne, Vic 3050, Australia
[3] Walter & Eliza Hall Inst Med Res, St Vincents Inst, Melbourne, Vic 3050, Australia
[4] Walter & Eliza Hall Inst Med Res, Bioinformat Div, Melbourne, Vic 3050, Australia