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Univ Kent, Inst Math Stat & Actuarial Sci, Canterbury CT2 7NF, Kent, England
Univ London Imperial Coll Sci Technol & Med, Fac Life Sci, Div Biol, Ascot SL5 7PY, Berks, England
Univ Cambridge, Dept Zool, Cambridge CB2 3EJ, England Univ Kent, Inst Math Stat & Actuarial Sci, Canterbury CT2 7NF, Kent, England