[1] Univ Manchester, Manchester Acad Hlth Sci Ctr, Ctr Genet & Genom Versus Arthrit, Manchester, Lancs, England
[2] Newcastle Univ, Newcastle Upon Tyne, England
[3] Musculoskeletal Directorate, Translat & Clin Res Inst, Newcastle Upon Tyne, England
[4] Univ Leeds, Leeds Inst Cardiovasc & Metab Med, Leeds, England
[5] Leeds Teaching Hosp Teaching Trust, NIHR Leeds BRC, Leeds, England
[6] Manchester Univ NHS Fdn Trust, NIHR Manchester Musculoskeletal Biomed Res Unit, Manchester, England
[7] Univ Manchester, Manchester Acad Hlth Sci Ctr, Versus Arthrit Ctr Epidemiol, Manchester, England
[8] Univ Coll Dublin, Sch Med & Med Sci, Dublin, Ireland