[1] DIPEQ, Inst Pesquisa Jardim Bot Rio de Janeiro, Rua Pacheco Leao 915, BR-22460030 Rio De Janeiro, RJ, Brazil
[2] Univ Fed Rio de Janeiro, Dept Bot, Museu Nacl, BR-20940040 Rio De Janeiro, RJ, Brazil
[3] Univ Nacl Mayor San Marcos, Fac Ciencias Biol, Posgrad Bot Trop, Av Venezuela,Cdra 34 S-N, Lima, Peru
[4] Herbario SP, Inst Bot, CP 68048, BR-04045972 Sao Paulo, SP, Brazil