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Northwest Univ, Coll Life Sci, Key Lab Resource Biol & Biotechnol Western China, Shaanxi Key Lab Anim Conservat, 229 North Taibai Rd, Xian 710069, Shaanxi, Peoples R China Nat Biodiversiot Ctr, Dept Terr Zool, Postbus 9517, NL-2300 RA Leiden, Netherlands