[1] Univ La Frontera, Butamallin Res Ctr Global Change, Av Francisco Salazar 01145, Temuco 4780000, Chile
[2] Univ La Frontera, Fac Agr & Forest Sci, Dept Forest Sci, Av Francisco Salazar 01145, Temuco 4780000, Chile
[3] Chinese Acad Sci, Inst Mt Hazards & Environm, Key Lab Mt Hazards & Earth Surface Proc, Chengdu 610041, Peoples R China