[1] Univ Liverpool, Inst Integrat Biol, Liverpool, Merseyside, England
[2] Univ Amsterdam, Inst Biodivers & Ecosyst Dynam, Amsterdam, Netherlands
[3] Queen Mary Univ London, Sch Biol & Chem Sci, London, England
[4] Nord Univ, Fac Biosci & Aquaculture, Bodo, Norway