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Chinese Acad Sci, CAS Key Lab Trop Marine Bioresources & Ecol, Guangdong Key Lab Marine Mat Med,South China Sea, Inst South China Sea Ecol & Environm Engn,RNAM Ct, Guangzhou 510301, Peoples R China Chinese Acad Sci, CAS Key Lab Trop Marine Bioresources & Ecol, Guangdong Key Lab Marine Mat Med,South China Sea, Inst South China Sea Ecol & Environm Engn,RNAM Ct, Guangzhou 510301, Peoples R China