Monophyly and phylogenetic placement of the spider genus Labulla Simon, 1884 (Araneae, Linyphiidae) and description of the new genus Pecado

The genus Labulla Simon is circumscribed in phylogenetic terms to include the species Labulla thoracica (Wider), L. flahaulti Simon and L. machadoi sp. nov. The genital anatomy of the genus is described in detail and the taxonomy of the genus is reviewed. The monophyly of Labulla is supported by numerous morphological apomorphies of the male palp and female epigynum. Based on a cladistic analysis, a new genus, Pecado gen. nov., is erected to place Labulla impudica Denis, from Northern Africa. Lepthyphantes insularis Saito and 'Labulla' nepula Tikader, both formerly included in Labulla, are not congeneric with the type species of Labulla. (c) 2005 The Linnean Society of London.