Review of Dissomphalus Ashmead, 1893 (Hymenoptera, Bethylidae) from Espirito Santo, Brazil, with description of twenty-one new species

Forty-two species of Dissomphalus Ashmead, 1893 are reviewed, 21 of them are described and illustrated: Dissomphalus botocudus sp. nov., D. fredi sp. nov., D. tupinikim sp. nov., D. guarani sp. nov., D. congo sp. nov., D. rosangelae sp. nov., D. w-aedeagus sp. nov., D. amana sp. nov., D. potyra sp. nov., D. pyata sp. nov., D. clovisi sp. nov., D. kuara sp. nov., D. miriamae sp. nov., D. cacirus sp. nov., D. mirim sp. nov., D. secretus sp. nov., D. caparao sp. nov., D. capixaba sp. nov., D. ibirapitanga sp. nov., D. purius sp. nov. and D. taiabocu sp. nov. Six species are registered for the first time from Espirito Santo: D. brasiliensis Kieffer, D. bahiensis Redighieri & Azevedo, D. dumosus Evans, D. incomptus Evans, D. refertus Alencar & Azevedo and D. umbilicus Azevedo. Three species were reidentified: D. napo Evans, D. truncatus-Azevedo and D. vallensis Azevedo. Among fifteen species-groups which recognized in the genus, amana species-group; amplus species-group and secretus species-group are newly proposed.