[1] Guangdong Ocean Univ, Dept Marine Technol, Zhanjiang, Peoples R China
[2] Southern Marine Sci & Engn Guangdong Lab Zhuhai, Zhuhai, Peoples R China
[3] Nanjing Univ Informat Sci & Technol, Sch Marine Sci, Nanjing, Peoples R China
[4] Zhanjiang Bay Lab Guangdong Prov, Zhanjiang, Peoples R China
[5] Univ Calif Los Angeles, Dept Atmospher & Ocean Sci, Los Angeles, CA 90095 USA