[1] Fisheries Res Agcy, Natl Res Inst Fisheries Sci, Aquat Genom Res Ctr, Kanazawa Ku, Kanagawa 2368648, Japan
[2] Fisheries Res Agcy, Res Ctr Fisheries & Environm Ariake & Yatsushiro, Seikai Natl Fisheries Res Inst, Nagasaki 8512213, Japan
[3] Nihon Univ, Lab Fish Pathol, Dept Vet Med, Kanagawa 2520880, Japan