[1] Univ Fed Parana, Programa Posgrad Bot, Caixa Postal 19031, BR-81531970 Curitiba, PR, Brazil
[2] CORBIDI, Div Ecol Vegetal, Calle Santa Rita 105 2, Lima, Peru
[3] Herbario Selva Cent Oxapampa, HOXA Estn Biol Jardin Botanico Missouri, Prolongacion Bolognesi Mz E-6, Pasco, Peru
[4] CORBIDI, Div Mastozool, Calle Santa Rita 105 2, Lima, Peru
[5] Louisiana State Univ, Dept Biol Sci, Shirley C Tucker Herbarium, Baton Rouge, LA 70803 USA
[6] Univ Nacl Mayor San Marcos, Museo Hist Nat, Dept Ecol, Av Arenales 1256, Lima, Peru
[7] New York Bot Garden, Bronx, NY 10445 USA