[1] Kobe Univ, Sch Med, Dept Anat & Brain Sci, Kobe, Hyogo 6500017, Japan
[2] Tohoku Inst Technol, Dept Elect, Sendai, Miyagi 9828577, Japan
[3] Erasmus Univ, Dept Cell Biol & Genet, MGC, NL-3000 DR Rotterdam, Netherlands
[4] Kumamoto Univ, Sch Med, Ctr Anim Resources & Dev, Inst Mol Embryol & Genet, Kumamoto 8620976, Japan
[5] Waseda Univ, Sch Human Sci, Dept Pharmacol & Brain Sci, Tokorozawa, Saitama 3591192, Japan