[1] Univ Westminster, Sch Biosci, London, England
[2] Royal Free & UCL Med Sch, London, England
[3] Kingston Univ, Sch Life Sci, Kingston upon Thames KT1 2EE, Surrey, England
[4] Brest Univ, Brest, France
[5] Univ Southampton, Tenovus Res Labs, Southampton SO9 5NH, Hants, England
[6] Univ Washington, Seattle, WA 98195 USA