[1] South China Normal Univ, Sch Geog, Guangzhou 510631, Guangdong, Peoples R China
[2] Southern Marine Sci & Engn Guangdong Lab, Zhuhai 519082, Peoples R China
[3] Chinese Acad Sci, Synth Res Ctr Chinese Ecosyst Res Network, Key Lab Ecosyst Network Observat & Modeling, Inst Geog Sci & Nat Resources Res, Beijing 100101, Peoples R China
[4] Univ Chinese Acad Sci, Coll Resources & Environm, Beijing 100190, Peoples R China