[1] INP ENSAT, EcoLab, Lab Ecol Fonct, F-31326 Castanet Tolosan, France
[2] CNRS INP UPS, EcoLab, Lab Ecol Fonct, UMR 5245, F-31326 Castanet Tolosan, France
[3] ISA, LGCgE, Equipe Sols & Environm, F-59046 Lille, France
[4] Univ Toulouse, Toulouse, France
[5] Univ Toulouse 3, F-31062 Toulouse, France