[1] Yale Univ, Sch Med, Dept Cell Biol, New Haven, CT 06510 USA
[2] Yale Univ, Sch Med, Dept Psychiat, New Haven, CT 06510 USA
[3] Yale Univ, Sch Med, Howard Hughes Med Inst, New Haven, CT 06510 USA
[4] Rockefeller Univ, Mol & Cellular Neurosci Lab, New York, NY 10021 USA