[1] Michigan State Univ, Dept Plant Biol, E Lansing, MI 48824 USA
[2] Smithsonian Inst, Natl Museum Nat Hist, Dept Bot, Washington, DC 20013 USA
[3] Chinese Acad Sci, Inst Bot, State Key Lab Vegetat & Environm Change, Beijing 100093, Peoples R China
[4] Smithsonian Conservat Biol Inst, Ecol Grp, Front Royal, VA 22630 USA
[5] Univ Puerto Rico, Inst Trop Ecosyst Studies, San Juan, PR 00936 USA
[6] Chinese Acad Sci, Key Lab Plant Resource Conservat & Sustainable Ut, S China Bot Garden, Guangzhou 510650, Guangdong, Peoples R China
[7] Univ Wisconsin, Dept Nat & Appl Sci, Green Bay, WI 54311 USA
[8] Chinese Acad Forestry, Res Inst Trop Forestry, Guangzhou 510520, Guangdong, Peoples R China
[9] Ctr Ecol & Hydrol Edinburgh, Penicuik EH26 0QB, Midlothian, Scotland
[10] Columbia Univ, Dept Ecol Evolut & Environm Biol, New York, NY 10027 USA
[11] Smithsonian Trop Res Inst, Balboa Ancon, Panama