[1] IDR Inst Desenvolvimento Rural Parana, IAPAR, EMATER, Rodovia Celso Garcia Cid,Km 375, BR-86047902 Londrina, PR, Brazil
[2] CIRAD, UMR AGAP, F-34398 Montpellier, France
[3] Univ Montpellier, CIRAD, INRA, Montpellier SupAgro, Montpellier, France
[4] CIRAD, UMR IPME, F-34398 Montpellier, France
[5] EMBRAPA Cafe Empresa Brasileira Pesquisa Agr, BR-70770901 Brasilia, DF, Brazil