[1] Stellenbosch Univ, Dept Bot & Zool, Ctr Invas Biol, Private Bag X1, ZA-7602 Stellenbosch, South Africa
[2] Nurtingen Geislingen Univ Appl Sci HFWU, Schelmenwasen 4-8, D-72622 Nurtingen, Germany
[3] Kirstenbosch Res Ctr, South African Natl Biodivers Inst, Private Bag X7, ZA-7735 Cape Town, South Africa
[4] Univ Toronto, Dept Ecol & Evolutionary Biol, 25 Willcocks St, Toronto, ON, Canada
[5] Univ Toronto Scarborough, Dept Biol Sci, 1265 Mil Trail, Toronto, ON, Canada
[6] Univ Fed Lavras, Dept Biol, Lavras, Brazil