[1] Hebei Univ, Coll Life Sci, Baoding 071002, Hebei, Peoples R China
[2] Chinese Acad Sci, Inst Bot, State Key Lab Vegetat & Environm Change, Beijing 100093, Peoples R China
[3] Peking Univ, Sino French Inst Earth Syst Sci, Coll Urban & Environm Sci, Beijing 100871, Peoples R China
[4] UMRCEA CNRS, LSCE, CE, Batiment 709, F-91191 Gif Sur Yvette, France
[5] Chinese Acad Sci, Inst Appl Ecol, Shenyang 110016, Liaoning, Peoples R China
[6] Chinese Acad Sci, Northwest Inst Plateau Biol, Xining 810008, Qinghai, Peoples R China