[1] UFZ Helmholtz Ctr Environm Res, Dept Isotope Biogeochem, D-04318 Leipzig, Germany
[2] Univ Grenoble Alpes, Inst Biol Struct, CNRS, CEA, 71 Ave Martyrs, F-38044 Grenoble, France
[3] Univ Grenoble Alpes, Lab Chim & Biol Met, UMR 5249, CNRS,CEA, 17 Ave Martyrs, F-38054 Grenoble, France
[4] ESRF, 71 Ave Martyrs, F-38043 Grenoble, France
[5] Royal Netherlands Inst Sea Res, NIOZ, Dept Marine Microbiol & Biogeochem, POB 59, NL-1790 AB Den Burg, Netherlands
[6] Univ Utrecht, POB 59, NL-1790 AB Den Burg, Netherlands
[7] UPMC Univ Paris 06, Sorbonne Univ, CNRS, Lab Oceanog Villefranche,UMR7093,Observ Oceanol, F-06 Villefranche Sur Mer, France
[8] Univ Grenoble Alpes, Cell & Plant Physiol Lab, CNRS, CEA,INRA, F-38054 Grenoble 9, France
[9] Univ Grenoble Alpes, CEA, Inst Nanosci & Cryogenie, F-38054 Grenoble, France
[10] European Mol Biol Lab, Cell Biol & Biophys Unit, D-69117 Heidelberg, Germany