[1] Univ Liverpool, Inst Syst Mol & Integrat Biol, Dept Mol & Clin Canc Med, Sherrington Bldg,Ashton St, Liverpool L69 3BX, Merseyside, England
[2] Clatterbridge Canc Ctr, Dept Phys, Wirral, Merseyside, England
[3] Univ Liverpool, Dept Phys, Oliver Lodge Lab, Liverpool, Merseyside, England
[4] Bhabha Atom Res Ctr, Radiol Phys & Advisory Div, Mumbai, Maharashtra, India
[5] PTW Freiburg, Freiburg, Germany
[6] Univ Manchester, Fac Biol Med & Hlth, Sch Med Sci, Div Canc Sci, Manchester, Lancs, England
[7] Glan Clwyd Gen Hosp, North Wales Canc Treatment Ctr, Radiotherapy Phys Dept, Bodelwyddan LL18 5UJ, Wales