[1] Arthritis Res UK Tissue Engn Ctr, Kobe, Hyogo, Japan
[2] Univ York, Dept Biol, York YO10 5DD, N Yorkshire, England
[3] Newcastle Univ, Musculoskeletal Res Grp, Inst Cellular Med, Newcastle Upon Tyne NE1 7RU, Tyne & Wear, England
[4] Newcastle Upon Tyne Hosp NHS Fdn Trust, Newcastle Upon Tyne NE1 7RU, Tyne & Wear, England
[5] Univ Aberdeen, Musculoskeletal Programme, Sch Med & Dent, Inst Med Sci, Aberdeen AB25 2ZD, Scotland