[1] Operat Wallacea Ltd, Wallace House, Spilsby PE23 4EX, Lincs, England
[2] Trinity Coll Dublin, Sch Nat Sci, Dept Zool, Dublin D02 CX56, Ireland
[3] Univ Coll Dublin, Sch Biol & Environm Sci, Dublin D04 N2E5, Ireland
[4] Univ Halu Oleo, Dept Biol & Biotechnol, South East Sulawesi, Indonesia