[1] Equilibrium Res, 47 Quays,Cumberland Rd, Bristol BS1 6UQ, Avon, England
[2] Univ Queensland, Sch Earth & Environm Sci, Brisbane, Qld 4072, Australia
[3] Open Univ, Dept Geog, Milton Keynes MK7 6AA, Bucks, England
[4] Open Univ, Open Space Res Ctr, Milton Keynes MK7 6AA, Bucks, England
[5] Cranfield Univ, Sch Water Energy & Environm, Coll Rd, Cranfield MK43 0AL, Beds, England
[6] Int Union Conservat Nat, Rue Mauverney 28, CH-1196 Gland, Switzerland
[7] ESiLi, Het Haam 16, NL-6846 KW Arnhem, Netherlands
[8] Chicago Bot Garden, 1000 Lake Cook Rd, Glencoe, IL 60022 USA
[9] 5 Marshall Rd, Cambridge CB1 7TY, England
[10] UCL, Dept Anthropol, Taviton St, London WC1H 0BW, England