[1] Gunma Univ, Adv Sci Res Leaders Dev Unit, Maebashi, Gunma, Japan
[2] Gunma Univ, Inst Mol & Cellular Regulat, Metab Signal Res Ctr, Maebashi, Gunma, Japan
[3] Kyushu Univ, Res & Dev Ctr Taste & Odor Sensing, Div Sensory Physiol, Fukuoka, Japan
[4] Monell Chem Senses Ctr, 3500 Market St, Philadelphia, PA 19104 USA
[5] Gunma Univ, Inst Mol & Cellular Regulat, Dept Cell Biol, Maebashi, Gunma, Japan
[6] Jichi Med Univ, Sch Med, Dept Physiol, Div Integrat Physiol, Shimotsuke, Japan