[1] Acad Sci Czech Republ, Inst Parasitol, CR-37005 Ceske Budejovice, Czech Republic
[2] Fisheries Agcy Japan, Natl Res Inst Far Seas Fisheries, Shimizu, Shizuoka 424, Japan
[3] Univ Tokyo, Dept Aquat Biosci, Grad Sch Agr & Life Sci, Bunkyo Ku, Tokyo 1138657, Japan