[1] Beijing Normal Univ, State Key Lab Earth Surface Proc & Resource Ecol, Beijing 100875, Peoples R China
[2] Beijing Normal Univ, Sch Geog, Beijing 100875, Peoples R China
[3] Nagoya Univ, Grad Sch Environm Studies, Nagoya, Aichi, Japan
[4] USDA ARS, Grazinglands Res Lab, El Reno, OK USA