[1] Univ Johannesburg, Dept Zool, Auckland Pk Campus, ZA-2006 Johannesburg, South Africa
[2] Univ Cantabria, Fac Med, Dept Anat & Cell Biol, Santander 39011, Spain
[3] Univ Messina, Polo Univ Annunziata, Dept Vet Sci, I-98168 Messina, Italy
[4] Univ Messina, Dept Chem Biol Pharmaceut & Environm Sci, Messina, Italy
[5] Nord Univ, Fac Biosci & Aquaculture, Bodo, Norway
[6] Natl Res Council CNR, Inst Marine Biol Resources & Biotechnol IRBIM, Sect Messina, Messina, Italy