[1] Hokkaido Univ, Grad Sch Agr, Dept Bot & Agron, Lab Crop Sci, Sapporo, Hokkaido 0608589, Japan
[2] Tohoku Agr Res Ctr, Natl Agr & Food Res Org NARO, Morioka, Iwate 0200198, Japan
[3] Indian Council Agr Res, Directorate Onion & Garl Res, Pune 410505, Maharashtra, India