[1] Dalian Nat Hist Museum, Dalian, Liaoning, Peoples R China
[2] Fed Univ ABC, Lab Paleontol Vertebrados & Comportamento Anim, Sao Paulo, Brazil
[3] Pavol Jozef Safarik Univ, Ctr Interdisciplinary Biosci Technol & Innovat Pk, PaleoBioImaging Lab, Evolutionary Biodivers Res Grp, Kosice, Slovakia
[4] Hainan Trop Ocean Univ, Sanya, Hainan, Peoples R China
[5] Chinese Acad Geol Sci, Inst Geol, Beijing, Peoples R China
[6] Hokkaido Univ, Sch Sci, Sapporo, Hokkaido, Japan
[7] Hokkaido Univ, Hokkaido Univ Museum, Sapporo, Hokkaido, Japan
[8] Beipiao Pterosaur Museum China, Beipiao, Liaoning, Peoples R China