[1] ENIS, Lab Eau Energie & Environm, Sfax, Tunisia
[2] ENIS, LRAE, Sfax, Tunisia
[3] Univ Aberdeen, Dept Plant & Soil Sci, Inst Biol & Environm Sci, Aberdeen AB24 3UU, Scotland
[4] Univ Neuchatel, Lab Soil Biol, CH-2009 Neuchatel, Switzerland
[5] ENIS, Lab Genie Enzymat & Microbiol, Sfax, Tunisia