[1] Humboldt Univ, Geog Dept, Unter den Linden 6, D-10099 Berlin, Germany
[2] Univ Nacl Tucuman, Inst Ecol Reg, CONICET, 4107 Yerba Buena, San Miguel De Tucuman, Argentina
[3] Humboldt Univ, Integrat Res Inst Transformat Human Environm Syst, Unter den Linden 6, D-10099 Berlin, Germany
[4] IRB, Inst Recursos Biol, Inst Nacl Tecnol Agr INTA, CIRN, De los Reseros & Las Cabanas S-N HB1712WAA, Hurlingham, Buenos Aires, Argentina