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Univ Fed Rio de Janeiro, Inst Biol, Dept Zool, CCS,Ilha Fundao, Bloco A,Class A0-108, BR-2240090 Rio De Janeiro, RJ, Brazil Univ Fed Rio de Janeiro, Inst Biol, Dept Zool, CCS,Ilha Fundao, Bloco A,Class A0-108, BR-2240090 Rio De Janeiro, RJ, Brazil