[1] IFREMER, CNRS, UM2, UMR MARBEC 248,IRD, Ave Jean Monnet,CS 30171, F-34203 Sete, France
[2] Univ Cape Town, Dept Biol Sci, Private Bag X3, ZA-7701 Cape Town, South Africa
[3] Univ La Rochelle, CNRS, UMR7372, Ctr Etud Biol Chize, F-79360 Villiers En Bois, France
[4] UBO, CNRS, UMR LEMAR 195, IRD, Campus Ifremer,BP 70, F-29280 Plouzane, France
[5] Univ Reunion, UMR ENTROPIE, CNRS, IRD, Ave Rene Cassin CS92003, F-97744 St Denis 9, Reunion, France
[6] Univ Toulon & Var, Aix Marseille Univ, CNRS, IRD,MIO, Marseille, France