[1] Chinese Acad Sci, Inst Tibetan Plateau Res, Key Lab Tibetan Environm Changes & Land Surface P, Beijing 100101, Peoples R China
[2] Yunnan Univ, Inst Palaeontol, Yunnan Key Lab Palaeobiol, Kunming 650500, Yunnan, Peoples R China
[3] Yunnan Univ, MEC Int Joint Lab Palaeobiol & Palaeoenvironm, Kunming 650500, Yunnan, Peoples R China
[4] Lake Biwa Museum, 1091 Oroshimo, Kusatsu, Shiga 5250001, Japan
[5] Shantou Univ, Inst Marine Sci, Shantou 515063, Peoples R China