[1] Univ Tokyo, Inst Mol & Cellular Biosci, Yayoi, Tokyo, Japan
[2] KIRIN, Pharmaceut Res Lab, Takasaki, Gumma, Japan
[3] Univ Tsukuba, Inst Basic Med Sci, Tsukuba, Ibaraki, Japan
[4] Univ Tsukuba, Ctr Tsukuba Adv Res Alliance, Tsukuba, Ibaraki, Japan