[1] Kindai Univ, Fac Agr, Dept Environm Management, 3327-204 Nakamachi, Nara 6318505, Japan
[2] Natl Univ Singapore, Fac Sci, Lee Kong Chian Nat Hist Museum, Singapore 117377, Singapore
[3] Indonesian Inst Sci, Res Ctr Oceanog, Bitung Marine Life Conservat Ctr, Jl Tandurusa, Bitung 95527, North Sulawesi, Indonesia