[1] Univ Ulsan, Dept Biochem & Mol Biol, Coll Med, Seoul 05505, South Korea
[2] Univ Ulsan, Asan Med Ctr, Dept Gastroenterol, Coll Med, Seoul 05505, South Korea
[3] Seoul Natl Univ, Dept Biomed Sci, Coll Med, Seoul 03080, South Korea
[4] Genome Inst Singapore, Human Genet Grp, Singapore, Singapore