[1] Univ Helsinki, Helsinki, Finland
[2] Helsinki Univ Hosp, Dept Med, Helsinki, Finland
[3] Minerva Fdn, Helsinki, Finland
[4] Univ Helsinki, Inst Mol Med Finland, FIMM, Helsinki, Finland
[5] Helsinki Univ Hosp, Dept Gastrointestinal Surg, Abdominal Ctr, Helsinki, Finland
[6] Helsinki Univ Hosp, Dept Pathol, Helsinki, Finland
[7] Orebro Univ, Sch Med Sci, Orebro, Sweden
[8] Univ Turku, Turku, Finland
[9] Abo Akad Univ, Turku Biosci Ctr, Turku, Finland
[10] Orebro Univ, Sch Sci & Technol, Orebro, Sweden