[1] Shaanxi Normal Univ, Sch Geog & Tourism, West Changan St 620, Xian 710119, Shaanxi, Peoples R China
[2] Chinese Acad Sci, Northwest Inst Ecoenvironm & Resources, State Key Lab Cryospher Sci, Lanzhou 730000, Peoples R China
[3] Indiana Univ Purdue Univ Indianapolis IUPUI, Dept Earth Sci, Indianapolis, IN 46202 USA
[4] Qinling Natl Bot Garden, Xian 710061, Peoples R China