[1] Inst Reg Arides, Lab Aridoculture & Cultures Oasiennes, Medenine 4119, Tunisia
[2] Agronutrition, Equipe Rech Agron, F-31390 Carbonne, France
[3] Univ Toulouse 3, Univ Toulouse, Lab Interact Mol & React Chim & Photochim, UMR CNRS 5623, F-31062 Toulouse, France