[1] UCL, Res Dept Infect & Populat Hlth, Mortimer Market Ctr, Capper St, London WC1E 6JB, England
[2] St Georges Univ London, Inst Infect & Immun, London, England
[3] Queen Mary Univ London, Blizard Inst, Ctr Immunol & Infect Dis, Barts & London Sch Med & Dent, London, England
[4] Brunel Univ London, Coll Engn Design & Phys Sci, London, England
[5] St Georges Univ London, Populat Hlth Res Inst, London, England