[1] Lanzhou Univ, Coll Pastoral Agr Sci & Technol, State Key Lab Grassland Agroecosyst, Lanzhou 730020, Peoples R China
[2] Hebei Univ Environm Engn, Hebei Key Lab Agroecological Safety, Qinhuangdao 066102, Hebei, Peoples R China
[3] Chengdu Univ Informat Technol, Resources & Environm Coll, Chengdu 610225, Peoples R China
[4] Chinese Acad Sci, Northwest Inst Plateau Biol, Key Lab Cold Reg Restorat, Xining 810008, Peoples R China
[5] Qinghai Normal Univ, Dept Life Sci, Xining 810008, Peoples R China